[May one] doubt whether, in cheese and timber, worms are generated, or, if beetles and wasps, in cow-dung, or if butterflies, locusts, shellfish, snails, eels, and such life be procreated of putrefied matter, which is to receive the form of that creature to which it is by formative power disposed[?] To question this is to question reason, sense, and experience. If he doubts this, let him go to Egypt, and there he will find the fields swarming with mice begot of the mud of the Nylus, to the great calamity of the inhabitants. A seventeenth century opinion quoted by L. L. Woodruff, in *The Evolution of Earth and Man*, 1929
I heard the mice too, rattling behind the panels, as if the same occurrence were important to their interests. But the black beetles took no notice of the agitation, and groped about the hearth in a ponderous elderly way, as if they were short-sighted and hard of hearing, and not on terms with one another.
The candles that lighted that room of hers were placed in sconces on the wall. They were high from the ground, and they burnt with the steady dulness of artificial light in air that is seldom renewed. As I looked round at them, and at the pale gloom they made, and at the stopped clock, and at the withered articles of bridal dress upon the table and the ground, and at her own awful figure with its ghostly reflection thrown large by the fire upon the ceiling and the wall, I saw in everything the construction that my mind had come to, repeated and thrown back to me. My thoughts passed into the great room across the landing where the table was spread, and I saw it written, as it were, in the falls of the cobwebs from the centre-piece, in the crawlings of the spiders on the cloth, in the tracks of the mice as they betook their little quickened hearts behind the panels, and in the gropings and pausings of the beetles on the floor.
Then, I looked round and saw the disturbed beetles and spiders running away over the floor, and the servants coming in with breathless cries at the door. I still held her forcibly down with all my strength, like a prisoner who might escape; and I doubt if I even knew who she was, or why we had struggled, or that she had been in flames, or that the flames were out, until I saw the patches of tinder that had been her garments no longer alight but falling in a black shower around us.
He ordered his horse to be saddled and, leaving his regiment on the march, rode to his father's estate where he had been born and spent his childhood. Riding past the pond where there used always to be dozens of women chattering as they rinsed their linen or beat it with wooden beetles, Prince Andrew noticed that there was not a soul about and that the little washing wharf, torn from its place and half submerged, was floating on its side in the middle of the pond. He rode to the keeper's lodge. No one at the stone entrance gates of the drive and the door stood open. Grass had already begun to grow on the garden paths, and horses and calves were straying in the English park. Prince Andrew rode up to the hothouse; some of the glass panes were broken, and of the trees in tubs some were overturned and others dried up. He called for Taras the gardener, but no one replied. Having gone round the corner of the hothouse to the ornamental garden, he saw that the carved garden fence was broken and branches of the plum trees had been torn off with the fruit. An old peasant whom Prince Andrew in his childhood had often seen at the gate was sitting on a green garden seat, plaiting a bast shoe.
It was of no avail that the pavements of Paris were there on every side, the classic and splendid hotels of the Rue de Varennes a couple of paces away, the dome of the Invalides close at hand, the Chamber of Deputies not far off; the carriages of the Rue de Bourgogne and of the Rue Saint-Dominique rumbled luxuriously, in vain, in the vicinity, in vain did the yellow, brown, white, and red omnibuses cross each other's course at the neighboring cross-roads; the Rue Plumet was the desert; and the death of the former proprietors, the revolution which had passed over it, the crumbling away of ancient fortunes, absence, forgetfulness, forty years of abandonment and widowhood, had sufficed to restore to this privileged spot ferns, mulleins, hemlock, yarrow, tall weeds, great crimped plants, with large leaves of pale green cloth, lizards, beetles, uneasy and rapid insects; to cause to spring forth from the depths of the earth and to reappear between those four walls a certain indescribable and savage grandeur; and for nature, which disconcerts the petty arrangements of man, and which sheds herself always thoroughly where she diffuses herself at all, in the ant as well as in the eagle, to blossom out in a petty little Parisian garden with as much rude force and majesty as in a virgin forest of the New World.
"Um! I love you even without the brandy, but with scoundrels I am a scoundrel. Ivan is not going to Tchermashnya--why is that? He wants to spy how much I give Grushenka if she comes. They are all scoundrels! But I don't recognize Ivan, I don't know him at all. Where does he come from? He is not one of us in soul. As though I'd leave him anything! I shan't leave a will at all, you may as well know. And I'll crush Mitya like a beetle. I squash black-beetles at night with my slipper; they squelch when you tread on them. And your Mitya will squelch too. _Your_ Mitya, for you love him. Yes, you love him and I am not afraid of your loving him. But if Ivan loved him I should be afraid for myself at his loving him. But Ivan loves nobody. Ivan is not one of us. People like Ivan are not our sort, my boy. They are like a cloud of dust. When the wind blows, the dust will be gone.... I had a silly idea in my head when I told you to come to-day; I wanted to find out from you about Mitya. If I were to hand him over a thousand or maybe two now, would the beggarly wretch agree to take himself off altogether for five years or, better still, thirty-five, and without Grushenka, and give her up once for all, eh?"
Aristotle's term was adopted by Linnaeus (1758), and has been universally used by zoologists. The identification of the elytra of beetles with the fore-wings of other insects has indeed been questioned (1880) by F. Meinert, who endeavoured to compare them with the tegulae of Hymenoptera, but the older view was securely established by the demonstration in pupal elytra by J. G. Needham (1898) and W. L. Tower (1903), of nervures similar to those of the hind-wing, and by the proof that the small membranous structures present beneath the elytra of certain beetles, believed by Meinert to represent the whole of the true fore-wings, are in reality only the alulae. Entry: COLEOPTERA
These firm fore-wings, or elytra (fig. 1, A), are usually convex above, with straight hind margins (_dorsa_); when the elytra are closed, the two hind margins come together along the mid-dorsal line of the body, forming a _suture_. In many beetles the hind-wings are reduced to mere vestiges useless for flight, or are altogether absent, and in such cases the two elytra are often fused together at the suture; thus organs originally intended for flight have been transformed into an armour-like covering for the beetle's hind-body. In correlation with their heavy build and the frequent loss of the power of flight, many beetles are terrestrial rather than aerial in habit, though a large proportion of the order can fly well. Entry: COLEOPTERA
ADEPHAGA.--This tribe includes beetles of carnivorous habit with five segments on every foot, simple thread-like feelers with none of the segments enlarged to form club or pectination, and the outer lobs (galea) of the first maxilla usually two-segmented and palpiform (fig. 4 b). The transverse fold of the hind-wing is towards the tip, about two-thirds of the wing-length from the base. At this fold the median nervure stops and is joined by a cross nervure to the radial, which can be distinguished throughout its length from the subcostal. There are four malpighian tubules. In the ovarian tubes of Adephaga small yolk-chambers alternate with the egg-chambers, while in all other beetles there is only a single large yolk-chamber at the narrow end of the tube. The larvae (fig. 2 c) are active, with well-chitinized cuticle, often with elongate tail-feelers (cerci), and with five-segmented legs, the foot-segment carrying two claws. Entry: ADEPHAGA
Students of the Coleoptera have failed to agree not only on a system of classification, but on the relative specialization of some of the groups which they all recognize as natural. Lameere, for example, considers some of his _Cantharidiformia_ as the most primitive Coleoptera. J. L. Leconte and G. H. Horn placed the _Rhynchophora_ (weevils) in a group distinct from all other beetles, on account of their supposed primitive nature. Kolbe, on the other hand, insists that the weevils are the most modified of all beetles, being highly specialized as regards their adult structure, and developing from legless maggots exceedingly different from the adult; he regards the Adephaga, with their active armoured larvae with two foot-claws, as the most primitive group of beetles, and there can be little doubt that the likeness between larvae and adult may safely be accepted as a primitive character among insects. In the Coleoptera we have to do with an ancient yet dominant order, in which there is hardly a family that does not show specialization in some point of structure or life-history. Hence it is impossible to form a satisfactory linear series. Entry: FIG
The _Curculionidae_, or weevils (q.v.), comprising 23,000 species, are by far the largest family of the group. The maxillary palps are short and rigid, and there is no distinct labrum, while the feelers are usually of an "elbowed" form, the basal segment being very elongate (figs. 41, 42). They are vegetable feeders, both in the perfect and larval stages, and are often highly injurious. The female uses her snout as a boring instrument to prepare a suitable place for egg-laying. The larvae (fig. 3) of some weevils live in seeds; others devour roots, while the parent-beetles eat leaves; others, again, are found in wood or under bark. The _Scolytidae_, or bark-beetles, are a family of some 1500 species, closely allied to the _Curculionidae_, differing only in the feeble development of the snout. They have clubbed feelers, and their cylindrical bodies (fig. 43) are well adapted for their burrowing habits under the bark of trees. Usually the mother-beetle makes a fairly straight tunnel along which, at short intervals, she lays her eggs. The grubs, when hatched, start galleries nearly at right angles to this, and when fully grown form oval cells in which they pupate; from these the young beetles emerge by making circular holes directly outward through the bark. Entry: RHYNCHOPHORA
The _Bruchidae_, or seed-beetles, agree with the two preceding families in tarsal structure; the head is largely hidden by the pronotum, and the elytra are short enough to leave the end of the abdomen exposed (fig. 38). The development of the pea and bean-beetles has been carefully studied by C. V. Riley, who finds that the young larva, hatched from the egg laid on the pod, has three pairs of legs, and that these are lost after the moult that occurs when the grub has bored its way into the seed. In Great Britain the beetle, after completing its development, winters in the seed, waiting to emerge and lay its eggs on the blossom in the ensuing spring. Entry: ANCHISTOPODA
The _Lampyridae_ are a large family, of which the glow-worm (_Lampyris_) and the "soldier beetles" (_Telephorus_) are familiar examples. The female "glow-worm" (fig. 15, b), emitting the well-known light (see above), is wingless and like a larva; the luminosity seems to be an attraction to the male, whose eyes are often exceptionally well developed. Some male members of the family have remarkably complex feelers. In many genera of _Lampyridae_ the female can fly as well as the male; among these are the South European "fireflies." Entry: MALACODERMATA
Locusts have many enemies besides man. Many birds greedily devour them, and it has many times been remarked that migratory swarms of the insects were closely followed by myriads of birds. Predatory insects of other orders also attack them, especially when they are in the unwinged condition. Moreover, they have still more deadly insect foes as parasites. Some attack the fully developed winged insect. But the greater part attack the eggs. To such belong certain beetles, chiefly of the family _Cantharidae_, and especially certain two-winged flies of the family _Bombyliidae_. These latter, both in the Old and New World, must prevent vast quantities of eggs from producing larvae. Entry: A
In all climates fruit and forest trees suffer from weevils or _Curculionidae_. The plum curculio (_Conotrachelus nenuphar_, Herbst) in America causes endless harm in plum orchards; curculios in Australia ravage the vines and fruit trees (_Orthorrhinus klugii_, Schon, and _Leptops hopei_, Bohm, &c.). In Europe a number of "long-snouted" beetles, such as the raspberry weevils (_Otiorhynchus picipes_), the apple blossom weevil (_Anthonomus pomorum_), attack fruit; others, as the "corn weevils" (_Calandra oryzae_ and _C. granaria_), attack stored rice and corn; while others produce swollen patches on roots (_Ceutorhynchus sulcicollis_), &c. All these _Curculionidae_ are very timid creatures, falling to the ground at the least shock. This habit can be used as a means of killing them, by placing boards or sacks covered with tar below the trees, which are then gently shaken. As many of these beetles are nocturnal, this trapping should take place at night. Larval "weevils" mostly feed on the roots of plants, but some, such as the nut weevil (_Balaninus nucum_), live as larvae inside fruit. Seeds of various plants are also attacked by weevils of the family _Bruchidae_, especially beans and peas. These seed-feeders may be killed in the seeds by subjecting them to the fumes of bisulphide of carbon. The corn weevils (_Calandra granaria_ and _C. oryzae_) are now found all over the world, in many cases rendering whole cargoes of corn useless. Entry: ECONOMIC
A very large number of other insects belonging to the same family possess blistering properties, owing to their containing cantharidin. Of these the most remarkable is the Telini "fly" of India (_Mylabris cichorii_), the range of which extends from Italy and Greece through Egypt and central Asia as far as China. It is very rich in cantharidin, yielding fully twice as much as ordinary cantharides. Several green-coloured beetles are, on account of their colour, used as adulterants to cantharides, but they are very easily detected by examination with the eye, or, if powdered, with the microscope. Entry: A
HEXAPODA (Gr. [Greek: hex], six, and [Greek: pous], foot), a term used in systematic zoology for that class of the ARTHROPODA, popularly known as insects. Linnaeus in his _Systema naturae_ (1735) grouped under the class Insecta all segmented animals with firm exoskeleton and jointed limbs--that is to say, the insects, centipedes, millipedes, crustaceans, spiders, scorpions and their allies. This assemblage is now generally regarded as a great division (phylum or sub-phylum) of the animal kingdom and known by K. T. E. von Siebold's (1848) name of Arthropoda. For the class of the true insects included in this phylum, Linnaeus's old term Insecta, first used in a restricted sense by M. J. Brisson (1756), is still adopted by many zoologists, while others prefer the name Hexapoda, first used systematically in its modern sense by P. A. Latreille in 1825 (_Familles naturelles du règne animal_), since it has the advantage of expressing, in a single word, an important characteristic of the group. The terms "Hexapoda" and "hexapod" had already been used by F. Willughby, J. Ray and others in the late 17th century to include the active larvae of beetles, as well as bugs, lice, fleas and other insects with undeveloped wings. Entry: HEXAPODA
_Forms of Larva._--The unlikeness of the young insect to its parent is one of the factors that necessitates metamorphosis. It is instructive, further, to trace among metabolic insects an increase in the degree of this dissimilarity. An adult Hexapod is provided with a firm, well-chitinized cuticle and six conspicuous jointed legs. Many larval Hexapods might be defined in similar general terms, unlike as they are to their parents in most points of detail. Examples of such are to be seen in the grubs of may-flies, dragon-flies, lacewing-flies and ground-beetles (fig. 24). This type of active, armoured larva--often bearing conspicuous feelers on the head and long jointed cercopods on the tenth abdominal segment--was styled campodeiform by F. Brauer (1869), on account of its likeness in shape to the bristle-tail _Campodea_. As an extreme contrast to this campodeiform type, we take the maggot of the house-fly (fig. 25)--a vermiform larva, with soft, white, feebly-chitinized cuticle and without either head-capsule or legs. Between these two extremes, numerous intermediate forms can be traced: the grub (wireworm) of a click-beetle, with narrow elongate well-armoured body, but with the legs very short; the grub of a chafer, with the legs fairly developed, but with the cuticle of all the trunk-segments soft and feebly chitinized; the well-known caterpillar of a moth (fig. 21, e) or saw-fly, with its long cylindrical body, bearing the six shortened thoracic legs and a variable number of pairs of "pro-legs" on the abdomen (this being the eruciform type of larva); the soft, white, wood-boring grub of a longhorn-beetle or of the saw-fly _Sirex_, with its stumpy vestiges of thoracic legs; the large-headed but entirely legless, fleshy grub of a weevil; and the legless larva, with greatly reduced head, of a bee. The various larvae of the above series, however, have all a distinct head-capsule, which is altogether wanting in the degraded fly maggot. These differences in larval form depend in part on the surroundings among which the larva finds itself after hatching; the active, armoured grub has to seek food for itself and to fight its own battles, while the soft, defenceless maggot is provided with abundant nourishment. But in general we find that elaboration of imaginal structure is associated with degradation in the nature of the larva, eruciform and vermiform larvae being characteristic of the highest orders of the Hexapoda, so that unlikeness between parent and offspring has increased with the evolution of the class. Entry: FIG
The importance of great cardinal features of the life-history as indicative of relationship leads us to consider the Endopterygota as a natural assemblage of orders. The occurrence of weevils--among the most specialized of the Coleoptera--in Triassic rocks shows us that this great order of metabolous insects had become differentiated into its leading families at the dawn of the Mesozoic era, and that we must go far back into the Palaeozoic for the origin of the Endopterygota. In this view we are confirmed by the impossibility of deriving the Endopterygota from any living order of Exopterygota. We conclude, therefore, that the primitive stock of the former sub-class became early differentiated from that of the latter. So widely have most of the higher orders of the Hexapoda now diverged from each other, that it is exceedingly difficult in most cases to trace their relationships with any confidence. The Neuroptera, with their similar fore- and hind-wings and their campodeiform larvae, seem to stand nearest to the presumed isopteroid ancestry, but the imago and larva are often specialized. The campodeiform larvae of many Coleoptera are indeed far more primitive than the neuropteran larvae, and suggest to us that the Coleoptera--modified as their wing-structure has become--arose very early from the primitive metabolous stock. The antiquity of the Coleoptera is further shown by the great diversity of larval form and habit that has arisen in the order, and the proof afforded by the hypermetamorphic beetles that the campodeiform preceded the eruciform larva has already been emphasized. Entry: RELATIONSHIPS
The prothorax is large and "free," i.e. readily movable on the mesothorax, an arrangement usual among insects with the power of rapid running. The tergite of the prothorax (pronotum) is prominent in all beetles, reaching back to the bases of the elytra and forming a substantial shield for the front part of the body. The tergal regions of the mesothorax and of the metathorax are hidden under the pronotum and the elytra when the latter are closed, except that the mesothoracic scutellum is often visible--a small triangular or semicircular plate between the bases of the elytra (fig. 1, A). The ventral region of the thoracic skeleton is complex, each segment usually possessing a median sternum with paired episterna (in front) and epimera (behind). The articular surfaces of the haunches (coxae) of the fore-legs are often conical or globular, so that each limb works in a ball-and-socket joint, while the hind haunches are large, displacing the ventral sclerites of the first two abdominal segments (fig. 1, C). The legs themselves (fig. 1, A) are of the usual insectan type, but in many families one, two, or even three of the five foot-segments may be reduced or absent. In beetles of aquatic habit the intermediate and hind legs are modified as swimming-organs (fig. 2, a), while in many beetles that burrow into the earth or climb about on trees the fore-legs are broadened and strengthened for digging, or lengthened and modified for clinging to branches. The hard fore-wings (elytra) are strengthened with marginal ridges, usually inflected ventrally to form epipleura which fit accurately along the edges of the abdomen. The upper surface of the elytron is sharply folded inwards at intervals, so as to give rise to a regular series of external longitudinal furrows (striae) and to form a set of supports between the two chitinous layers forming the elytron. The upper surface often shows a number of impressed dots (punctures). Along the sutural border of the elytron, the chitinous lamella forms a tubular space within which are numerous glands. The glands occur in groups, and lead into common ducts which open in several series along the suture. Sometimes the glands are found beneath the disk of the elytron, opening by pores on the surface. The hind-wings, when developed, are characteristic in form, possessing a sub-costal nervure with which the reduced radial nervure usually becomes associated. There are several curved median and cubital nervures and a single anal, but few cross nervures or areolets. The wing, when not in use, is folded both lengthwise and transversely, and doubled up beneath the elytron; to permit the transverse folding, the longitudinal nervures are interrupted. Entry: COLEOPTERA
EARWIG, an insect belonging to the _Forficulidae_, a family usually referred to the Orthoptera, but sometimes regarded as typifying a special order, to which the names Dermaptera, Dermatoptera and Euplexoptera have been given, in allusion to certain peculiarities in the structure of the wings in the species that possess them. The front wings are short and horny and when at rest meet without overlapping in the middle line, like the wing-cases of brachelytrous (cocktail) beetles. The hind wings, on the contrary, are for the most part membranous and, when extended, of large size; each consists of two portions, the distal of which, in virtue of the arrangement and jointing of its nervures, is capable of being both doubled up and folded fanwise beneath the proximal, which is partly horny when the wing is tucked away under the front wing-case of the same side. Apart from these characteristics, the most distinctive feature of earwigs is the presence at the end of the abdomen of a pair of pincers which are in reality modified appendages, known as cercopods, and represent the similar limbs of _Japyx_ and the caudal feelers of _Campodea_ and some other insects. Entry: EARWIG